Lucy Kerhoulas

Associate Professor - Forest Ecophysiology

I study ecophysiology in coniferous forests. My research investigates the relationships among water use, growth, physiology, climate, and forest management. I use stable isotope analyses, dendrochronology, and gas exchange measurements as my primary research tools. Learn more about my research by visiting the Forest Physiology Lab.


Ph.D. Northern Arizona University, Biology
M.S. Humboldt State University, Biology
B.S. Humboldt State University, Botany

Courses Taught

FOR 130 - Dendrology
FOR 331 - Silvics: The Foundation of Silviculture

Current Graduate Students

Name Thesis
Gabriel Roletti
Sophia Lemmo


Beckmann, J.J., R.L. Sherriff, L.P. Kerhoulas, and J.M. Kane. 2021. Douglas-fir encroachment reduces drought resistance in Oregon white oak of northern California. Forest Ecology and Management 498: 119543. (PDF)

Gorman, A.J., L.P. Kerhoulas, W.T. Polda, and N.J. Kerhoulas. 2019. Epiphyte diversity and distribution in an old Sitka spruce crown. Evansia 36: 12
22. (PDF) 

Kerhoulas, L.P., and J.M. Kane. 2011. Sensitivity of ring growth and carbon allocation to climatic variation vary within ponderosa pine trees. Tree Physiology 32: 14–23. (PDF)

Kerhoulas, L.P., D.T. Hammons, and N.J. Kerhoulas. 2020. Bigleaf maple within-crown leaf morphology and seasonal physiology. Northwest Science 94: 176-187. (PDF)

Kerhoulas, L.P., T.E. Kolb, and G.W. Koch. 2017. The influence of monsoon climate on latewood growth in southwestern ponderosa pine. Forests 8: 116. (PDF)

Kerhoulas, L.P., T.E Kolb, and G.W. Koch. 2013. Tree size, stand density, and the source of water used across seasons by ponderosa pine in northern Arizona. Forest Ecology and Management 289: 425–433. (PDF)

Kerhoulas, L.P., W.T. Polda, N.J. Kerhoulas, and J.-P. Berrill. 2020. Physiology and growth of Douglas-fir and redwood seedlings planted after partial harvesting. Frontiers in Forests and Global Change 3: 49. (PDF)

Kerhoulas, L.P., A.S. Weisgrau, E.C. Hoeft, and N.J. Kerhoulas. 2020. Vertical gradients in foliar physiology of tall Picea sitchensis trees. Tree Physiology: 1-12. (PDF)

Kerhoulas, L.P., T.E Kolb, M.D. Hurteau and G.W. Koch. 2013. Managing climate change adaptation in forests: a case study from the U.S. Southwest. Journal of Applied Ecology 50: 1311–1320. (PDF)

Mullin, L.P., Sillett, S.C., Koch, G.W., K.P. Tu, and M.E. Antoine. 2009. Physiological consequences of height-related morphological variation in Sequoia sempervirens foliage. Tree Physiology 29: 999–1010. (PDF)

Soland, K.R., L.P. Kerhoulas, N.J. Kerhoulas, and J.R. Teraoka. 2021. Second-growth redwood forest responses to restoration treatments. Forest Ecology and Management 496: 119370. (PDF)

van Mantgem, P.J., L.P. Kerhoulas, R.L. Sherriff, and Z.J. Wenderott. 2020. Tree-ring evidence of forest management moderating drought responses: implications for dry, coniferous forests in the southwestern U.S. Frontiers in Forests and Global Change 3: 41. (PDF)

Weinberg, W.C., J.R. Suoja, L.P. Kerhoulas, R.J. Maberry, C.A. Lee, D.S. Baston, and S.E. Marshall. 2021. Phytophthora ramorum foliar infection reduces leaf-level productivity in tanoak and California by: A pilot study from Redwood National Park. Madroño 68: 99-108. (PDF)
Lucy Kerhoulas
(707) 826-4243
115 Natural Resources Building